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Published articles - Behaviour and welfare - general



Abbott, B. B., Schoen, L. S. & Badia, P. (1984). Predictable and unpredictable shock: behavioral measures of aversion and physiological measures of stress. Psychological Bulletin, 96 (1), 45.Link


Badia, P., Culbertson, S. & Harsh, J. (1973). Choice of longer or stronger signalled shock over shorter or weaker unsignalled shock. Journal of the Experimental Analysis of Behavior, 19 (1), 25-32. Link


Badia, P., Harsh, J. & Abbott, B. (1979). Choosing between predictable and unpredict- able shock conditions: Data and theory. Psychological Bulletin, 86 (5), 1107-1131.Link


Barnett, J. & Hemsworth, P. (1990). The validity of physiological and behavioural measures of animal welfare. Applied Animal Behaviour Science,  25, 177-187.Link


Bassett, L. & Buchanan-Smith, H. M. (2007). Effects of predictability on the welfare of captive animals. Applied Animal Behaviour Science, 102 (3-4), 223-245. Link


Boissy, A., Manteuffel, G., Jensen, M. B., Moe, R. O., Spruijt, B., Keeling, L. J., Winckler, C., Forkman, B., Dimitrov, I., Langbein, J. et al.  (2007).  Assessment of positive emotions in animals to improve their welfare. Physiology & Behavior,  92 (3), 375–397. Link


Broom, D. (1986). Indicators of poor welfare. British Veterinary Journal, 142 (6), 524-526.Link


Broom, D. (1991a). Animal welfare: concepts and measurement. Journal of Animal Science, 69 (10), 4167-4175.Link


Broom, D. (1991b). Assessing welfare and suffering. Behavioural Processes, 25 (2), 117–123.


Broom, D. (2006). Behaviour and welfare in relation to pathology. Applied Animal Behaviour Science, 97 (1), 73-83.Link


Broom, D. & Johnson, K. (1993). Stress and animal welfare. Dordrecht: Kluwer Chapman and Hall.Link


Broom, D. & Kennedy, M. (1993). Stereotypies in horses: their relevance to welfare and causation. Equine Veterinary Education, 5 (3), 151-154.Link


Broom, D. & Kirkden, R. (2004). Welfare, stress, behaviour and pathophysiology. Veterinary Pathophysiology, 337-369.Link


Burman, O., Parker, R. M. A., Paul,  E. S. & Mendl, M.  (2008).  Sensitivity to reward loss as an indicator of animal emotion and welfare. Biology Letters, 4 (4), 330-333.Link


Byrne, A. & Eysenck, M. W. (1993). Individual differences in positive and negative interpretive biases. Personality and Individual Differences, 14 (6), 849–851.Link

Dawkins, M. (2004). Using behaviour to assess animal welfare. Animal Welfare, 13 (SUPPL.), S3-S7.Link


Dawkins, M. (2006). A user’s guide to animal welfare science. Trends in Ecology & Evolution, 21 (2), 77–82.Link


Dawkins, M. (2013). The science of animal welfare. London: Springer. Link


DeVries, A. C., Glasper, E. R. & Detillion, C. E. (2003). Social modulation of stress responses. Physiology & Behavior, 79 (3), 399–407.Link


Fraser,  D.  (2008).  Understanding  animal  welfare.  Acta  Veterinaria Scandinavica, 50 (SUPPL. 1).Link


Hansen, S. W. & Jeppesen, L. L. (2006). Temperament, stereotypies and anticipatory behaviour as measures of welfare in mink. Applied Animal Behaviour Science, 99 (1), 172-182.Link


Harding, E. J., Paul, E. S. & Mendl, M. (2004). Animal behaviour: cognitive bias and affective state. Nature, 427 (6972), 312-312.Link



Lyons, D. M., Parker, K. J., Katz, M. & Schatzberg, A. F. (2009). Developmental cascades linking stress inoculation, arousal regulation, and resilience. Frontiers in Behavioral Neuroscience, 3, 32.Link


Koolhaas, J., Korte, S., De Boer, S., Van  Der Vegt,  B., Van  Reenen, C., Hopster, H.,   De Jong, I., Ruis, M. & Blokhuis, H. (1999). Coping styles in animals: current status in behavior and stress-physiology. Neuroscience & Biobehavioral Reviews, 23 (7), 925-935.Link


Martin P & Bateson P 2007. Measuring Behaviour: an Introductory Guide. Cambridge University Press, Cambridge, UK.Link


Mason, G. & Latham, N. (2004). Can’t stop, won’t stop: Is stereotypy a reliable animal welfare indicator? Animal Welfare, 13 (SUPPL.), S57-S69.Link


Mendl, M., Burman, O., Parker, R. & Paul, E.  (2009).  Cognitive bias as an indicator of animal emotion and welfare: Emerging evidence and underlying mechanisms. Applied Animal Behaviour Science, 118 (3-4), 161-181.Link


Mendl, M., Burman,  O. & Paul,  E.  (2010).  An integrative and functional framework  for the study of animal emotion and mood. Proceedings of the Royal Society B: Biological Sciences, 277 (1696), 2895-2904.Link



Panksepp, J. (2011).  The basic emotional circuits of mammalian brains:  Do animals  have affective lives? Neuroscience & Biobehavioral Reviews, 35 (9), 1791-1804.Link


Poole, T. (1992). The nature and evolution of behavioural needs in mammals. Animal Welfare, 1 (3), 203–220.Link


Rogers, L. J. (2010). Relevance of brain and behavioural lateralization to animal welfare. Applied Animal Behaviour Science, 1 127 (1), 1–11.Link


Vyas A, Pillai AG, Chattarji S (2004) Recovery after chronic stress fails to reverse amygdaloid neuronal hypertrophy and enhanced anxiety-like behavior. Neuroscience 128(4):667–3.Link


Wiepkema, P. & Koolhaas, J. (1993). Stress and animal welfare. Animal Welfare, 2 (3), 195-218.Link


Yeates, J. & Main, D. (2008). Assessment of positive welfare: a review. The Veterinary Journal, 175 (3), 293-300. Link